In classical antiquity, several theses were elaborated on the origin of the Etruscans from the 5th century BC, when the Etruscan civilization had been already established for several centuries in its territories, that can be summarized into three main hypotheses.
The first is the autochthonous development in situ out of the Villanovan culture, as claimed by the Greek historian Dionysius of Halicarnassus who described the Etruscans autochthonous people who had always lived in Etruria.Dionysius of Halicarnassus, Roman Antiquities, Book I Chapter 30 1.
The second is a migration from the Aegean Sea, as claimed by two Greek historians: Herodotus, who described them as a group of immigrants from Lydia in Anatolia, and Hellanicus of Lesbos who claimed that the Tyrrhenians were the Pelasgians originally from Thessaly, Greece, who entered Italy at the head of the Adriatic Sea in Northern Italy.Dionysius of Halicarnassus, Roman Antiquities, 1.17–19
The third hypothesis was reported by Livy and Pliny the Elder, and puts the Etruscans in the context of the Rhaetian people to the north and other populations living in the Alps.Titus Livius (Livy), The History of Rome (Ab Urbe Condita), Book 5
The first Greek author to mention the Etruscans, whom the Ancient Greeks called Tyrrhenians, was the 8th-century BC poet Hesiod, in his work, the Theogony. He mentioned them as residing in central Italy alongside the Latins.Hesiod, Theogony 1015. The 7th-century BC Homeric Hymn to DionysusHomeric Hymn to Dionysus, 7.7–8 referred to them as pirates.John Pairman Brown, Israel and Hellas, Vol. 2 (2000) p. 211 Unlike later Greek authors, such as Herodotus and Hellanicus, these earlier Greek authors did not suggest that Etruscans had migrated to Italy from elsewhere.
According to prehistoric and protohistoric archaeologists, anthropologists, etruscologists, geneticists, linguists, all the evidence gathered so far points to an autochthonous origin of the Etruscans.
Helmut Rix's classification of the Etruscan language within the Tyrsenian family—alongside Raetic and Lemnian—has gained support from comparative linguistics. While the discovery of Lemnian inscriptions in 1885 once suggested a possible east-to-west migration, more recent linguistic and archaeological assessments argue instead for a west-to-east diffusion. Scholars such as Wallace (2010), Simon (2021), and Chiai (2024) interpret the Lemnian language as a derivative or dialect of Etruscan introduced to the island through maritime contact, possibly via an Etruscan trading enclave, rather than evidence of a common origin or population movement, as argued previously by Gras, De Simone and Robert Drews.
A mtDNA study published in 2013 concluded that the Etruscans' mtDNA appears very similar to that of Neolithic Europe population from Central Europe and to other Tuscan populations. This coincides with the Rhaetic language, which was spoken south and north of the Alps in the area of the Urnfield culture of Central Europe. The Villanovan culture, the early period of the Etruscan civilization, derives from the Proto-Villanovan culture that branched from the Urnfield culture around 1200 BC. An autochthonous population that diverged genetically was previously suggested as a possibility by Cavalli-Sforza.Cavalli-Sforza, L. L., P. Menozzi, A. Piazza. 1994. The History and Geography of Human Genes. Princeton University Press, Princeton.
A 2019 genetic study published in the journal Science analyzed the autosomal DNA of 11 Iron Age samples from the areas around Rome, concluding that Etruscans (900-600 BC) and the Latins (900-200 BC) from Latium vetus were genetically similar, and Etruscans also had Steppe-related ancestry despite speaking a pre-Indo-European language.
A 2021 genetic study published in the journal Science Advances analyzed the autosomal DNA of 48 Iron Age individuals from Tuscany and Lazio and confirmed that the Etruscan individuals displayed the ancestral component Steppe in the same percentages as found in the previously analyzed Iron Age Latins, and that the Etruscans' DNA completely lacks a signal of recent admixture with Anatolia or the Eastern Mediterranean, concluding that the Etruscans were autochthonous and they had a genetic profile similar to their Latin neighbors. Both Etruscans and Latins joined firmly the European cluster, 75% of the Etruscan male individuals were found to belong to haplogroup R1b-M269 and its subclades, especially R1b-P312 and its derivative R1b-L2 whose direct ancestor is R1b-U152, while the most common mitochondrial DNA haplogroup among the Etruscans was H.
With this passage, Dionysius launched the autochthonous theory, that the core element of the Etruscans, who spoke the Etruscan language, were of "Terra (Earth) itself"; that is, on location for so long that they appeared to be the original or native inhabitants. They are therefore the owners of the Villanovan culture.Page 52. Pallottino attributes this theory in modern times to the historian, Eduard Meyer, with Ugo Antonielli later associating the Villanovan and the natives. But Mayer soon adopted the oriental theory and Antonielli the northern. Drews in The End of the Bronze Age, p. 59, available as a preview on Google Books at [1], reports on Meyer and the views of Antonielli are stated in a review by R. A. L. Fell of Studi Etruschi. Vol. I. Rassegna di Etruscologia by A. Neppi Modona, the first page of which is found at [2].
Picking up this theme, Bonfante (2002) states:Page 3.
An additional elaboration conjectures that the Etruscans werePallottino, page 52, who says that he relies on Alfredo Trombetti and Giacomo Devoto.
In 1942, the Italy History Massimo Pallottino published a book entitled The Etruscans (which would be released in English language in 1955). Pallottino presented various hypotheses that gained wide acceptance in the Archeology community. He said "no one would dream of asking where Italians or Frenchmen came from originally; it is the formation of the Italian and France nations that we study." He meant that the formation process for Etruscan civilization took place in Etruria or nearby. Formulating a different point of view on the same evidence, Pallottino says:
J. P. Mallory compares the Etruscans to other remnant non Indo-European central Mediterranean populations, such as the Basques of the Iberian Peninsula and southern France, who absorbed the art styles and alphabet of their Greek neighbors.Mallory (1989). In Search of the Indo-Europeans: Language, Archaeology and Myth. London: Thames & Hudson.
The British archaeologists, Graeme Barker and Tom Rasmussen, were also fervent supporters of the "autochthonous theory". In their book, The Etruscans, they state, "There is no evidence for the kind of cultural break at the Villanovan/Etruscan transition envisaged by either of the ‘plantation’ models from the eastern Mediterranean, or for a folk movement of either kind from continental Europe in the Late Bronze Age,".Barker, Graeme, and Tom Rasmussen. The Etruscans. Blackwell Publishers, 1998, pp. 44. Thus, inferring that the Etruscans were indigenous to Italy and descended from the previous communities of Etruria.
Many supporters of this theory also believed that the Etruscans had foreign influences on their culture. For instance, the historian, Mario Torelli agreed with Dionysius’s claims and believed that the Etruscans inherited elements of their culture from other Italic peoples.Torelli, Mario. “The Etruscan City-State.” A Comparative Study of Thirty City-State Cultures: An Investigation, 2000, pp. 192. Robert Leighton also agreed with the “autochthonous theory”, but he believed the Etruscan's culture was impacted by Greek and Phoenician merchants.Leighton, Robert. Tarquinia: An Etruscan City. Duckworth, 2004, pp. 44.
Since ancient times, doubts have been raised about the accuracy of Herodotus' claims. Xanthus of Lydia, originally from Sardis and a great connoisseur of the history of the Lydians, wasn't aware of a Lydian origin of the Etruscans, as reported by Greek historian Dionysius of Halicarnassus.
The classical scholar Michael Grant commented on this story, writing that it "is based on erroneous etymologies, like many other traditions about the origins of 'fringe' peoples of the Greek world". Grant writes there is evidence that the Etruscans themselves spread it to make their trading easier in Asia Minor when many cities in Asia Minor, and the Etruscans themselves, were at war with the Greeks.
The French scholar Dominique Briquel also disputed the historical validity of Herodotus' account. Briquel demonstrated that "the story of an exodus from Lydia to Italy was a deliberate political fabrication created in the Hellenized milieu of the court at Sardis in the early 6th century BC." Briquel also commented that "the traditions handed down from the Greek authors on the origins of the Etruscan people are only the expression of the image that Etruscans' allies or adversaries wanted to divulge. For no reason, stories of this kind should be considered historical documents".Dominique Briquel, Le origini degli Etruschi: una questione dibattuta sin dall’antichità, in M. Torelli (ed.), Gli Etruschi Catalogo, Bompiani, Milano, 2000, p. 43–51 (Italian).
However, the Greek History Dionysius of Halicarnassus objected that the Tyrrhenian (Etruscan) culture and language shared nothing with the Lydian. He stated:
During the 6th to 5th centuries BC, the word "Tyrrhenians" was referred to the Etruscans, for whom the Tyrrhenian Sea is named, according to Strabo. In Pindar, the Tyrsenoi appear grouped with the Carthaginians as a threat to Magna Graecia:
Thucydides mentions them together with the Pelasgians and associates them with Lemnos pirates and with the pre-Greek population of Attica. Lemnos remained relatively free of Greek influence up to Hellenistic times, and the Lemnos stele of the 6th century BC is inscribed with a language very similar to Etruscan. This has led to the postulation of a "Tyrrhenian language group" comprising Etruscan, Lemnian and Raetic language. There is thus linguistic evidence of a relationship between the Lemnians and the Etruscans. Some scholars ascribe this link to Etruscan expansion between the 8th and 6th centuries BC, putting the homeland of the Etruscans in Italy and the Alps particularly because of their relation to the Alpine Raetic population. Adherents of this latter school of thought point to the legend of Lydian origin of the Etruscans referred to by Herodotus, and the statement of Livy that the Raetians were Etruscans driven into the mountains by the invading Gauls. Critics of this theory point to the very scanty evidence of a linguistic relationship of Etruscan with Indo-European, let alone Anatolian in particular, and to Dionysius of Halicarnassus who decidedly argues against an Etruscan-Lydian relationship. The Indo-European Lydian language is first attested some time after the Tyrrhenian migrants are said to have left for Italy.
Classical authors such as Herodotus and Dionysius of Halicarnassus were among the first to offer explanations for Etruscan origins. Herodotus proposed a Lydian migration led by King Tyrrhenus, a theory consistent with the Greek tradition of explaining cultural origins through heroic migrations.Herodotus, Histories 1.94Larissa Bonfante, Etruscans: Life and Afterlife: A Handbook of Etruscan Studies, Wayne State University Press, 1986. Dionysius, by contrast, emphasized an autochthonous origin, suggesting the Etruscans were native to Italy and had no cultural or linguistic connection to Lydia.Dionysius of Halicarnassus, Roman Antiquities 1.30
Modern scholars tend to approach these ancient sources critically. John Bryan Ward-Perkins, for example, argued that Greek and Roman writers often projected political or cultural biases onto their accounts of other peoples. He referred to these narratives as “a hostile tradition” and described them as seen through “a veil of interpretation, misunderstanding, and at times, plain invention.”John Bryan Ward-Perkins, "The Problem of Etruscan Origins," Harvard University, 1959. Dominique Briquel similarly concluded that stories of eastern origin were often politically motivated and should not be regarded as historical evidence.
In the mid-20th century, Italian archaeologist Massimo Pallottino revived the autochthonous theory, arguing that Etruscan civilization emerged through a local process of cultural formation in Etruria. He acknowledged external influences but maintained that these occurred on Italian soil and did not constitute evidence of population replacement.Massimo Pallottino, The Etruscans, Indiana University Press, 1955.
Linguistic evidence has also shaped this debate. R.S.P. Beekes argued that the close relationship between the Etruscan and Lemnian languages supports a migration from the eastern Mediterranean.R.S.P. Beekes, The Origin of the Etruscans, Royal Dutch Academy, 2003. However, Alison E. Cooley countered that such similarities may result from cultural contact, such as trade, rather than shared ethnic origins.Alison E. Cooley, Review of R.S.P. Beekes, The Origin of the Etruscans, The Classical Association, 2005. Recent scholarship on the Raetic language and the hypothesized Tyrrhenian language family generally regards Lemnian and Etruscan as related languages. While the direction and nature of their relationship have been the subject of prolonged debate, an increasing number of scholars propose that the Lemnian language may reflect Etruscan influence transmitted through maritime interactions, resulting from a west-to-east movement.Rex E. Wallace, Zikh Rasna: A Manual of the Etruscan Language and Inscriptions, Beech Stave Press, 2010.Zsolt Simon, "The Alleged Anatolian Loanwords in Etruscan: A Reconsideration," in Federico Giusfredi and Zsolt Simon (eds.), Studies in the Languages and Language Contact in Pre-Hellenistic Anatolia (Barcino Monographica Orientalia 17), Universitat de Barcelona, 2021, pp. 227–242.Gian Franco Chiai, "Lemnos, its Culture and Cultural Memories," in Simona Marchesini (ed.), Rhaeti & Co.: Nuovi scenari sulla questione tirrenica, Mantova: SAP Società Archeologica, 2024.
Some modern sociolinguists emphasize the broader role of language in cultural identity formation. Kari Gibson has described language as a “powerful symbol of national and ethnic identity,” shaping how communities perceive themselves.Kari Gibson, "The Myths of Language Use and the Homogenization of Bilingual Workers’ Identities," University of Hawaii, 2004. Literary theorist Gloria Anzaldúa similarly described language and ethnicity as interwoven, stating: “Ethnic identity is twin skin to linguistic identity.”Gloria Anzaldúa, Borderlands/La Frontera: The New Mestiza, Aunt Lute Books, 1987, p. 59. While these perspectives are modern and do not directly address ancient societies, they are sometimes cited to highlight how language may contribute to group identity.
Archaeological evidence has provided critical insights into Etruscan society and its development. Burial sites, such as those in the Monterozzi necropolis in Tarquinia, offer rich material evidence. For example, the Tomb of the Leopards, dating to the 5th century BC, depicts men and women dining together with similar crowns, suggesting social roles for women that differ markedly from those in Greek society.Luisa Banti, Etruscan Cities and Their Culture, University of California Press, 1973. Pallottino argued that such representations reflect a distinct cultural worldview shaped within Etruria, rather than imported from abroad.
Other tombs, like the Tomb of Hunting and Fishing, depict scenes in which humans appear subordinate to nature—a reversal of the dominant artistic conventions in many ancient societies. Scholars interpret this as indicative of a local artistic and philosophical tradition.
Although Etruscan culture was influenced by interactions with Greeks, Phoenicians, and other Mediterranean peoples, modern scholars increasingly support the conclusion that the Etruscan civilization developed locally out of the Villanovan culture, with external contact shaping but not defining its formation.
Several archaeologists who have analyzed Bronze Age and Iron Age remains that were excavated in the territory of historical Etruria have pointed out that no evidence has been found, related either to material culture or to social practices, that can support a migration theory from the Aegean Sea. The most marked and radical change that has been archaeologically attested in the area is the adoption, starting in about the 12th century BC, of the funeral rite of incineration in terracotta urns, which is a Continental European practice, derived from the Urnfield culture; there is nothing about it that suggests an ethnic contribution from Asia Minor or the Near East. One of the most common mistakes for a long time, even among some scholars of the past, has been to associate the later Orientalizing period of Etruscan civilization, due, as has been amply demonstrated by archeologists, to contacts with the Greeks and the Eastern Mediterranean and not mass migrations, with the question of their origins. The facial features (the profile, almond-shaped eyes, large nose) in the frescoes and sculptures, and the depiction of reddish-brown men and light-skinned women, influenced by archaic Greek art, followed the artistic traditions from the Eastern Mediterranean, that had spread even among the Greeks themselves, and to a lesser extent also to other several civilizations in the central and western Mediterranean up to the Iberian Peninsula. Actually, many of the tombs of the Late Orientalizing and Archaic periods, such as the Tomb of the Augurs, the Tomb of the Triclinium or the Tomb of the Leopards, as well as other tombs from the archaic period in the Monterozzi necropolis in Tarquinia, were painted by Greek painters or, in any case, foreigner artists. These images have, therefore, a very limited value for a realistic representation of the Etruscan population. It was only from the end of the 4th century B.C. that evidence of physiognomic portraits began to be found in Etruscan art and Etruscan portraiture became more realistic.
A 2012 survey of the previous 30 years’ archaeological findings, based on excavations of the major Etruscan cities, showed a continuity of culture from the last phase of the Bronze Age (12th–10th century BC) to the Iron Age (9th–8th century BC). This is evidence that the Etruscan civilization, which emerged around 900 BC, was built by people whose ancestors had inhabited that region for at least the previous 200 years, as has also been confirmed by anthropological and genetic studies. Based on this cultural continuity, there is now a consensus among archeologists that Proto-Etruscan culture developed, during the last phase of the Bronze Age, from the indigenous Proto-Villanovan culture, and that the subsequent Iron Age Villanovan culture is most accurately described as an early phase of the Etruscan civilization. According to Renato Peroni, a prominent archaeologist specializing in the prehistory and protohistory of Italy and an advocate for the autochthonous origins of the Etruscans, the archaeological continuity in Etruria can be traced back at least to the Bell Beaker culture. A significant discontinuity is recorded only during the transition from the Chalcolithic to the Bronze Age (2300–1000 BC). From 2300 BC onward, Etruria exhibits substantial settlement continuity. During this period, a constellation of local groups, all broadly associated with the Bell Beaker tradition, is already discernible in Tuscany and adjacent regions.
It is possible that there were contacts between northern-central Italy and the Mycenaeans at the end of the Bronze Age. However, contacts between the inhabitants of Etruria and inhabitants of Greece, Aegean Sea Islands, Asia Minor, and the Near East are attested only centuries later, as well as those with the Celtic world, when Etruscan civilization was already flourishing and Etruscan ethnogenesis was well established. The first of these attested contacts relate to the Magna Grecia and the Nuragics and Sardo-Punics in Sardinia, and the consequent orientalizing period.
A 2019 genetic study published in the journal Science analyzed the autosomal DNA of 11 Iron Age samples from the areas around Rome, including for the first time the whole genome sequencing (WGS) of some samples from Etruscan tombs, and concluded that Etruscans (900-600 BC) and the Latins (900-200 BC) from Latium vetus were genetically similar, and Etruscans also had Steppe-related ancestry despite speaking a pre-Indo-European language. A 2021 genetic study published in the journal Science Advances analyzed the autosomal DNA of 48 Iron Age individuals from Tuscany and Lazio and confirmed that the Etruscan individuals displayed the ancestral component Steppe in the same percentages as found in the previously analyzed Iron Age Latins, and that the Etruscans' DNA completely lacks a signal of recent admixture with Anatolia or the Eastern Mediterranean, concluding that the Etruscans were autochthonous and they had a genetic profile similar to their Latin neighbors. Both Etruscans and Latins joined firmly the European cluster, 75% of the Etruscan male individuals were found to belong to haplogroup R1b-M269 and its subclades, especially R1b-P312 and its derivative R1b-L2 whose direct ancestor is R1b-U152, while the most common mitochondrial DNA haplogroup among the Etruscans was H. Iron Age Etruscans from central Italy could be modelled as deriving 50% of their ancestry from Central European Bell Beakers, represented by Germany Bell Beaker, with around 25-30% Steppe-related ancestry, and the rest of their ancestry from a local Chalcolithic population. The conclusions of these studies have been confirmed by later ones.
In the collective volume Etruscology published in 2017, British archeologist Phil Perkins provides an analysis of the state of DNA studies and writes that "none of the DNA studies to date conclusively prove that Etruscans were an intrusive population in Italy that originated in the Eastern Mediterranean or Anatolia" and "there are indications that the evidence of DNA can support the theory that Etruscan people are autochthonous in central Italy".
In his book A Short History of Humanity published in 2021, German geneticist Johannes Krause, co-director of the Max Planck Institute for Evolutionary Anthropology in Jena, concludes that it is likely that the Etruscan language (as well as Basque language, Paleo-Sardinian and Minoan language) "developed on the continent in the course of the Neolithic Europe".
The conclusions of the 2021 study are in line with those of an earlier 2019 study, the first to publish analyses of whole genome sequencing of Etruscan samples, although the 2019 is more focused on ancient Rome than the question of Etruscan origins. The 2019 genetic study, published in the journal Science, analyzed the remains of eleven Iron Age individuals from the areas around Rome, of which four were Etruscan individuals, one buried in Veio from the Villanovan period (900-800 BC) and three buried in La Mattonara Necropolis near Civitavecchia from the Orientalizing period (700-600 BC). The study concluded that Etruscans (900–600 BC) and the Latins (900–500 BC) from Latium vetus were genetically similar, genetic differences between the examined Etruscans and Latins were found to be insignificant. The Etruscan individuals and contemporary Latins were distinguished from preceding populations of Italy by the presence of 30.7% steppe ancestry. Their DNA was a mixture of two-thirds Copper Age ancestry (EEF + WHG; Etruscans ~66–72%, Latins ~62–75%) and one-third Steppe-related ancestry (Etruscans ~27–33%, Latins ~24–37%) (with the EEF component mainly deriving from Neolithic-era migrants to Europe from Anatolia and the WHG being local Western European hunter-gatherers, with both components, along with that from the steppe, being found in virtually all European populations). The only sample of Y-DNA extracted belonged to haplogroup J-M12 (J2b-L283), found in an individual dated 700-600 BC, and carried exactly the M314 derived allele also found in a Middle Bronze Age individual from Croatia (1631-1531 calBCE). While the four samples of mtDNA extracted belonged to haplogroups U5a1, H, T2b32, K1a4. Therefore, Etruscans had also Steppe-related ancestry despite speaking a pre-Indo-European language.
In 2024, 6 individuals of Etruscan remains from Tarquinia, Lazio, dated the 9th-7th Century BC, were studied and confirmed the previous finds, Etruscans were an indigenous population. The admixture model showed that they were 84-92% Italy Bell Beaker and 8-26% additional Yamnaya Samara (Steppe-related) ancestry, but with one individual being more similar to Iron Age populations from Scandinavia, and north-west Europe. The two male individuals studied for Y-Chromosome belonged to the J2b/J-M12 lineage, and the five studied mitochondrial haplogroups were typical of post-Neolithic Europe. Phenotypic traits showed blue-eyes, light/dark brown hair, and pale to intermediate skin tones.
A mtDNA study, published in 2018 in the journal American Journal of Physical Anthropology, compared both ancient and modern samples from Tuscany, from the Prehistory, Etruscan age, Ancient Rome, Renaissance, and Present-day, and concluded that the Etruscans appear as a local population, intermediate between the prehistoric and the other samples, placing in the temporal network between the Eneolithic and the Roman Age.
These results are largely in line with previous mtDNA results from 2004 (in a smaller study also based on ancient DNA), and contradictory to results from 2007 (based on modern DNA). The 2004 study was based on mitochondrial DNA (mtDNA) from 80 bone samples, reduced to 28 bone samples in the analysis phase, taken from dating from the seventh century to the third century BC from Veneto, Tuscany, Lazio and Campania. This study found that the ancient DNA extracted from the Etruscan remains had some affinities with modern European populations including Germans, English people, and Tuscans in Italy. The study was marred by concerns that mtDNA sequences from the archeological samples represented severely damaged or contaminated DNA; however, subsequent investigation showed that the samples passed the most stringent tests of DNA degradation available.
A recent Y-DNA study from 2018 on a modern sample of 113 individuals from Volterra, a town of Etruscan origin, Grugni at al. keeps all the possibilities open, although the autochthonous scenario is the most supported by numbers, and concludes that "the presence of J2a-M67* (2.7%) suggests contacts by sea with Anatolian people, the finding of the Central European lineage G2a-L497 (7.1%) at considerable frequency would rather support a Northern European origin of Etruscans, while the high incidence of European R1b lineages (R1b 49.8%, R1b-U152 24.5%) cannot rule out the scenario of an autochthonous process of formation of the Etruscan civilization from the preceding Villanovan society, as suggested by Dionysius of Halicarnassus". In Italy Y-DNA J2a-M67*, not yet found in Etruscan samples, is more widespread on the Adriatic Sea coast between Marche and Abruzzo, and not in those where once lived the Etruscans, and in the study has its peak in the Ionian side of Calabria. In 2014, a late Bronze Age Kyjatice culture sample in Hungary was found to be J2a1-M67, a couple of J2a1b were found in Late Neolithic samples from the LBK culture in Austria, a J2a1a was found in a Middle Neolithic Sopot culture sample from Croatia, a J2a was found in a Late Neolithic Lengyel culture sample from Hungary. In 2019, in a Stanford study published in Science, two ancient samples from the Neolithic settlement of Ripabianca di Monterado in the province of Ancona, in the Marche region of Italy, were found to be Y-DNA J-L26 and J-M304. In 2021, two more ancient samples from the Chalcolitich settlement of Grotta La Sassa, in the province of Latina in southern Lazio, were found to be Y-DNA J2a7-Z2397. Therefore, Y-DNA J2a-M67 may be likely in Italy since the Neolithic and can't be the proof of recent contacts with Anatolia. In any case, J2a-M67 was not found among the Etruscan samples, unlike G2a-L497 and R1b-U152 who were actually found in the Etruscan individuals in really significant percentages.
Recent studies on the population structure of modern-day Italians have shown that in Italy there is a north–south cline for Y-chromosome lineages and autosomal loci, with a clear differentiation of peninsular Italians from Sardinians, and that modern Tuscans are the population of central Italy closest genetically to the inhabitants of northern Italy. A 2019 study, based on autosomal DNA of 1616 individuals from all 20 Italian administrative regions, concludes that Tuscans join the northern Italian cluster, close to the inhabitants of Liguria and Emilia-Romagna. A 2013 study, based on uniparental markers of 884 unrelated individuals from 23 Italian locations, had shown that the structure observed for the paternal lineages in continental Italy and Sicily suggests a shared genetic background between people from Tuscany and Northern Italy from one side, and people from Southern Italy and the Adriatic coast from the other side. The most frequent Y-DNA haplogroups in the group represented by populations from North-Western Italy, including Tuscany and most of the Padana plain, are four R1b-lineages (R-U152*, R-M269*, R-P312* and R-L2*).
Historical claims of autochthonous (indigenous) origin
Historical claims of allochthonous (outside) origin
The Sea Peoples theory
Differentiating between cultural origin and cultural influence
Archeological evidence and modern etruscology
Genetic evidence
Archeogenetics and ancient DNA
Previous archaeogenetic studies on mtDNA only
Older studies based on modern samples only
Further reading
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